It might run on 3.1 and 3.11 via win32s, there were occasional odd things that would run on the win32 subsystem as was adapted to 3.1/3.11, but not 95/98/ME.
ryao's above proposition is the converse, that only Win32s in DOS+Windows ME was adequate to running this, and that the prior Win32s in DOS+Windows 98 and DOS+Windows 95 was not.
To be honest, though, I very much doubt that anyone has fully tested either that claim or what the headlined article itself claims. A quick look at the source turns up things that require the full Windows NT security model, wide character versions of the API, and threads.
It probably does not run on any version of DOS+Windows, and "except Win95 and Win98" in the headlined article is likely a fumbled way of saying that, which didn't account for those not being the only versions of DOS+Windows that exist.
> ryao's above proposition is the converse, that only Win32s in DOS+Windows ME was adequate to running this, and that the prior Win32s in DOS+Windows 98 and DOS+Windows 95 was not.
Win32s was for Windows 3.x only
Win32 for 9x/Me was originally called Win32c, but then Microsoft renamed it to just plain Win32, despite the fact that name was shared by the much more complete NT-based implementation.
It won't. I see lots of calls to CreateThread/etc in the source. Win32s didn't support threads.