[AnimalMuppet]

Well, all right, do ants form cities? Yes, they do, at least of a kind.

You could even argue that lichen is a city, inhabited by multiple species.

I mean, I'm not sure I'd go so far as to argue than either of those really are a civilization. But if "city" is your sole criterion...

[PaulHoule]

Or a beehive. Bees moved into the basement window of my house and watching them go in and out it seemed to me that this single beehive had more departures and arrivals than all the commercial airports in the world put together.

If you look at the problem of "bee decline" from the viewpoint of the beekeeper where you are responsible for it you are responsible for a "city" of 50,000 insects that faces all kinds of threats from the inside and outside.

[saalweachter]

Ant nests and beehives aren't cities, they are households, consisting of a single family.