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I think there are two axes: reason about and intuit. I "understand" ballistic flight when I can calculate a solution that puts an artillery round on target. I also "understand" ballistic flight when I make a free throw with a basketball.

On writing that, I have an instinct to revise it to move the locus of understanding in the first example to the people who calculated the ballistic tables, based on physics first-principles. That would be more accurate, but my mistake highlights something interesting: an artillery officer / spotter simultaneously uses both. Is theirs a "deeper" / "truer" understanding? I don't think it is. I don't know what I think that means, for humans or AI.