Not in my experience. The walls were dark. And the window was bright occupying half of the surface of one wall. They still flew in circles.
I think it might be what other commenters are pointing out. That flies consider the brightest direction to be the sky, but if it's to the side (with respect to gravity) it messes up their navigation.
I think it might be what other commenters are pointing out. That flies consider the brightest direction to be the sky, but if it's to the side (with respect to gravity) it messes up their navigation.