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Due to its historical influences, 8051-based microcontrollers and cores are still widely used in a huge number of embedded devices and ASICs everywhere - motherboard EC, Ethernet, Wi-Fi controllers, this also includes the peripheral devices that plug into USB ports, like a mouse, a keyboard, a sensor, or industrial controllers, or whatever, so the first part of the joke is most communications via USB are just 8051s talking to each others. The second part of the joke is that even USB ASICs themselves, like USB hubs or host controllers, are often powered by 8051 cores.