Interestingly, if you take a look at old screenshots of Gnome 2.x, GTK apps have had more "mac like" layout/distribution/whitespace of controls within application windows for a very long time, even if the Gnome itself took more after Win9x in other aspects (like having a taskbar).
By contrast KDE/Qt has always had a stronger Win9x feel through and through.
By contrast KDE/Qt has always had a stronger Win9x feel through and through.