Is there any video where the artist talks about the process? I'm really interested in understanding how the artist approached this. Anyways, beautiful work.
While there is some commit history, it's definitely not low level enough to understand the process fully.
But I am imagining, from a quick "inspect element" looking at a few of the elements, that it is inspired by "brush movements": basically, you do a single brush movement with a single styled/transformed element (probably even more granular since a brush would give you trail-lines by default).