[derefr]

On the other hand, organisms are generally very "fault tolerant" systems in the small—if you get a crucial gene copied, and then one of the copies mutates to do something weird while the other remains functioning, the resulting organism is probably going to be viable, since the weird thing that the mutated copy of the gene does will [unless it's some very specific type of "weird thing" it's producing] be treated by the body as just another foreign enzyme to xenometabolize or foreign antibody to attack.

And the organism's genetic line can persist with that mutant gene indefinitely, building onto it until it becomes a useful feature.

(This isn't so true in especially small organisms, where an extra gene here and there might blow their size or resource budget. This is one of several things constraining the evolution of mitochondria, for example.)