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I worked as an engineer at a startup early in my career (looong time ago). Company went under, stock => $0.

However, I am extremely grateful for that time. It taught me a wide range of non-technical skills that have proved invaluable throughout my career.

Most of those non-technical skills revolved around leadership opportunities, being given huge responsibilities vs. my skill/age (and having to figure it all out on my own), and getting to participate in many aspects of the business.

It ingrained in me ownership; and I've never been able to shake bringing that startup mentality to everything I do at a larger company.

Just my experience. Much of this perhaps is achievable while at larger companies, depending on the situation.