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There are a set of optimisations that you can only do at runtime that you can't do with AOT. Anything that depends on data is something that you can't reliably do, such as eliding null checks if you can prove this cannot happen based on the data passed in. There are also cases where you can have multiple subclasses of a type such as a normal and a debug subclass, or multiple drivers for different back ends such as MongoDB or Cassandra, only one of which is used at runtime but you cannot know ahead of time which is selected (for example, it's based on an environment variable or system property).

The point is that while AOT can do a set of optimisations, including whole module analysis, there are a set which are only available at runtime.