I think we're starting to see more of this sort of thing happening now with Proton and Wine gaining prominence in the Linux community. Some games (Elden Ring comes to mind) have bad enough PC ports when they come out that the compatibility layer can incorporate a hotfix to improve performance, while users of the software on the original platform still had to suffer.
People from Transmeta told me stories about how their translators were full of special case optimizations to fix horrors they discovered in Microsoft Windows itself.